Prof. Dr. Hartmut Arndt

Curriculum vitae

1975-1980 Study of biology at the University of Rostock (major fields: marine biology, limnology, fishery biology, animal physiology)
1980-1987 Graduate student and research assistant in the division of Experimental Ecology, Department of Biology, University of Rostock (Professors Schnese and Schiewer)
1985 Ph.D. thesis on population ecology of zooplankters in an inner coastal water of the Baltic at the University of Rostock
1987-1991 Research associate at the Department of Aquatic Ecosystems, Division of Hydrology (Academy of Sciences in Berlin) and Leipniz-Institute of Freshwater Ecology and Inland Fisheries
1990/1991 Fellowship of the Humboldt Foundation at the Max-Planck-Institute of Limnology (Plön, Germany) and at the Institute for Lake Research in Langenargen (Germany)
1991-1994 Research associate at the Institute of Limnology, Austrian Academy of Sciences, Mondsee, Austria
1993 Habilitation at the University of Rostock on the role of protozoans in microbial and macrobial food webs
1994-1997 Associate professor for ecology at the University of Greifswald, Institute of Ecology, Kloster /Hiddensee
since 1997 Full professor for animal ecology at the University of Cologne, Zoological Institute



Research interests

Experimental and theoretical ecology (esp. freshwater and marine organisms), complexity of ecosystems and chaos research, structure and function of pelagic and benthic microbial food webs (running waters, lakes, deep sea, polar regions, groundwater), trophic interactions in biofilm communities, global biodiversity and molecular ecology of protists (esp. heterotrophic flagellates ), physiological ecology of protists.


Recent publications


Borcherding, J., Kiel, S., Struebig, A., Arndt, H. (2017): Vertical migration of Daphnia galeata – Evidence for the use of an alternative resource from a lethal environment. Limnologica 62: 132-139

Jeuck, A., Nitsche, F., Wylezich, C., Wirth, O., Hennemann, M., Troll, N., Bergfeld, T., Monir, S., Scherwaß, A., Arndt, H. (in press) A comparison of quantification methods for heterotrophic flagellates of different taxonomic groups. Protist

Venter, P., Nitsche, F., Domonell, A., Heger, P., Arndt, H. (in press) The protistan microbiome of grassland soil: distribution in the mesoscale. Protist

Živaljić, S.,  Schoenle, A., Nitsche, F., Hohlfeld, M., Piechocki, J., Reif, F., Shumo, M., Weiss, A., Werner, J., Witt, M., Voss, J., Arndt, H. (in press) Survival of marine heterotrophic flagellates isolated from the surface and the deep sea at high hydrostatic pressure: literature review and own experiments. Deep-Sea Res. II


Arndt, H., Monsonís Nomdedeu, M. (2016) Protozoans and global climate change in aquatic systems, in: Marxsen, J. (Ed.), Climate Change and Microbial Ecology: Current Research and Future Trends. Caister Academic Press, Poole, UK, pp. 41-52

Borcherding, J., Arndt, H., Breiden, S., Brenner, K., Heermann, L., Höfer, S., Leistenschneider, C., Lindner, J., Staas, S., Gertzen, S. (2016): Drift of fish larvae and juveniles in the Lower Rhine before and after the goby invasion. Limnologica 59: 53–6

Gossner, M.M., Lewinsohn, T., Kahl, T., Grassein, F., Boch, S., Prati, D., Birkhofer, K., Renner, S.C., Sikorski, J., Arndt, H., Baumgartner, V., Blaser, S., Blüthgen, N., Börschig, C., Buscot, F., Diekötter, T., Jorge, L.R., Jung, K., Keyel, A.C., Klein, A.-M., Klemmer, S., Krauss, J., Lange, M., Müller, J., Overmann, J., Pašalić, E., Penone, C., Perović, D., Purschke, O., Schall, P., Socher, S.A., Sonnemann, I., Tschapka, M., Tscharntke, T., Türke, M., Venter, P.C., Weiner, C.N., Werner, M., Wolters, V., Wurst, S., Westphal, C., Wubet, T., Fischer, M, Weisser, W.W., Allan, E. (2016): Land-use intensification causes homogenization of grassland communities. Nature 540: 266-269

Groll, F., Arndt, H., Altland, A. (2016): Chaotic attractor in two-prey one-predator system originates from interplay of limit cycles. Theor Ecol: DOI: 10.1007/s12080-016-0317-9

Haileselasie, T.H., Mergeay, J., Weider, L.J., Sommaruga, R., Davidson, T.A., Meerhoff, M., Arndt, H., Jürgens, K., Jeppesen, E., De Meester, L. (2016): Environment not dispersal limitation drives clonal composition of arctic Daphnia in a recently deglaciated area. Molecular Ecology (in press)

Scherwass A., Erken M., Arndt H. (2016): Grazing effects of ciliates on microcolony formation in bacterial biofilms. In: "Microbial biofilms - Importance and Applications": 81-93 Ed: D. Dhanasekaran and N. Thajuddin

Schönle, A., Jeuck, A., Nitsche, F., Venter, P., Prausse, D., Arndt, H. (2016) Methodological studies on estimates of abundance and diversity of heterotrophic flagellates from the deep-sea floor. J. Mar. Sci. Eng. 4, 22; doi:10.3390/jmse4010022

Schössow, M., Arndt, H., Becker, G. (2016): Response of gastropod grazers of the Lower River Rhine to food conditions, current velocity, and substratum roughness. Limnologica 58: 49-58

Soliveres, S., Manning, P., Prati, D., Gossner, M.M., Alt, F., Arndt, H., Baumgartner, V., Binkenstein, J., Birkhofer, K., Blaser, S., Bluthgen, N., Boch, S., Bohm, S., Borschig, C.,  Buscot, F., Diekotter, T., Heinze, J., Holzel, J., Jung, K., Klaus, V.H., Klein, A.-M.,  Kleinebecker, T., Klemmer, S., Krauss, J., Lange, M., Morris, E.-K., Müller, J., Oelmann, Y., Overmann, J., Pašalić, E., Renner, S.C., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schoning, I., Schrumpf, M., Sikorski, J., Socher, S.A., Solly, E.F., Sonnemann, I., Sorkau, E., Steckel, J., Steffen-Dewenter, I., Stempfhuber, B.,  Tschapka, M., Turke, M., Venter, P., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Wolters, V., Wubet, T., Wurst, S., Fischer, M., Allan, E. (2016): Locally rare species influence grassland ecosystem multifunctionality. Phil. Trans. R. Soc. B 371: 20150269. doi:10.1098/rstb.2015.0269

Soliveres, S., van der Plas, F., Manning, P., Prati, D., Gossner, M.M., Renner, S.C., Alt, F., Arndt, H., Baumgartner, V., Binkenstein, J., Birkhofer, K., Blaser, S., Blüthgen, N., Boch, S., Böhm, S., Börschig, C., Buscot, F., Diekötter, T., Heinze, J., Hölzel, N., Jung, K., Klaus, V.H., Kleinebecker, T., Klemmer, S., Krauss, J., Lange, M., Morris, E.K., Müller, J., Oelmann, Y., Overmann, J., Pašalić, E., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schöning, I., Schrumpf, M., Sikorski, J., Socher, S.A., Solly, E.F., Sonnemann, I., Sorkau, E., Steckel, J., Steffan-Dewenter, I., Stempfhuber, B., Tschapka, M., Türke, M., Venter, V., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Wolters, V., Wubet, T., Wurst, S., Fischer, M., Allan, E. (2016): Biodiversity at multiple trophic levels is needed for ecosystem multifunctionality. Nature 536: 456–459

Weisse, T., Anderson, R., Arndt, H., Calbet, A., Hansen, P.J., Montagnes, D.J. (2016): Functional ecology of aquatic phagotrophic protists - concepts, limitations, and perspectives. Europ. J. Protistol. 55: 50–74


Nitsche, F., Arndt, H. (2015) Comparison of similar Arctic and Antarctic morphotypes of heterotrophic protists regarding its genotypes and ecotypes. Protist 166: 42-57

Schoolmann, G., Nitsche, F., Arndt, H. (2015) Aspects of the life span and phenology of the invasive freshwater shrimp Atyaephyra desmarestii (Millet, 1831) at the northeastern edge of its range (Upper Rhine). Crustaceana 88: 949-962


Deng, L., Krauss, S., Feichtmayer, J. Hofmann, R., Arndt, H., Griebler, C. (2014) Grazing of heterotrophic flagellates on viruses is driven by feeding behaviour. Environmental Microbiology Reports (2014) 6(4), 325–330,

Jeuck, A., Arndt, H., Nitsche, F. (2014): Extended phylogeny of the Craspedida (Choanomonada). Europ. J. Protistol. 50: 430-443


Becks, L., Arndt, H. (2013): Different types of synchrony in chaotic and cyclic communities. Nature Communications 4: 1359, 1-9, DOI:10.1038

Domonell, A., Brabender, M., Nitsche, F., Bonkowski, M., Arndt, H. (2013): Community structure of cultivable protists in different grassland and forest soils of Thuringia. Pedobiologia 56: 1-7

Jeuck, A., Arndt, H. (2013): A short guide to common heterotrophic flagellates of freshwater habitats based on the morphology of living organisms. Protist 164: 842-860

Quintela-Alonso, P., Nitsche, F., Wylezich, C., Arndt, H., Foissner, W. (2013): A New Tetrahymena (Ciliophora, Oligohymenophorea) from Groundwater of Cape Town, South Africa. J. Euk. Microbiol. 60:235-246


Arndt, H. (2012): Protozoen. In: Grundwasserbiologie - Grundlagen und Anwendungen. DVGW-Information Wasser Nr. 75: 70-78

Brabender, M., Kiss, A.K., Domonell, A., Nitsche, F., Arndt, H. (2012): Phylogenetic and morphological diversity of novel soil cercomonad species with a description of two new genera (Nucleocercomonas and Metabolomonas). Protist 163: 495-528

Erken, M., Farrenschon, N., Speckmann, S., Arndt, H., Weitere, M. (2012): Quantification of individual fagellate - bacteria interactions within semi-natural biofilms. Protist 163: 632-642

Monsonís Nomdedeu, M., Willen, C., Schieffer, A., Arndt, H. (2012): Temperature dependent ranges of coexistence in a model of a two-prey-one-predator microbial food web. Marine Biology 159:2423–2430

Salani, F.S., Arndt, H., Hausmann, K., Nitsche, F., Scheckenbach, F. (2012): Analysis of the community structure of abyssal kinetoplastids revealed similar communities at larger spatial scales. ISME Journal 6: 713-723

Stoupin, D., Kiss, A.K., Arndt, H., Shatilovich, A.V., Gilichinsky, D.A., Nitsche, F. (2012): Cryptic diversity within the choanoflagellate morphospecies complex Codosiga botrytis – Phylogeny and morphology of ancient and modern isolates. Europ. J. Protistol. 48: 263-273


Ackermann, B., Esser, M., Scherwaß, A., Arndt, H. (2011): Long-term dynamics of microbial biofilm communities of the River Rhine with special references to ciliates. Int. Revue Hydrobiol. 96:1-19

Bergfeld, T., Scherwass, S., Ackermann, B., Fischer, H., Arndt, H., Schöl, A. (2011): Longitudinal and seasonal dynamics of the planktonic microbial community along the length of the River Rhine. River Systems 19(4): 337-349

Howe, A.T., Bass, D., Scoble, J.M., Lewis, R., Vickerman, K., Arndt, H., Cavalier-Smith, T. (2011): Novel cultured protists identify deep-branching environmental DNA clades of Cercozoa: New genera Tremula, Micrometopion, Minimassisteria, Nudifila, Peregrinia. Protist 162: 332-372

Nitsche, F., Carr, M., Arndt, H., Leadbeater, B.S.C. (2011): Higher level taxonomy and molecular phylogenetics of the Choanoflagellatea. J. Eukaryot. Microbiol. 58: 452-462

Quintela Alonso, P., Nitsche, F., Arndt, H. (2011): Molecular Characterization and Revised Systematics of Microdiaphanosoma arcuatum (Ciliophora, Colpodea). J. Eukaryot. Microbiol. 58: 114-119


Norf, H., Kniggendorf, L.G., Fischer, A., Arndt, H., Kureck, A. (2010): Sexual and reproductive traits of (Polychaeta, Ampharetidae): a remarkable invasive species in Central European waterways. Freshw. Biol. 55, 2510-2520

Scheckenbach, F., Hausmann, K., Wylezich, C., Weitere, M., Arndt, H. (2010): Large-scale patterns in biodiversity of microbial eukaryotes from the abyssal sea floor. PNAS 107(1): 115-12

Scherwass, A., T. Bergfeld, A. Schoel, M. Weitere und H. Arndt (2010): Changes in the plankton community along the length of the River Rhine: Lagrangian sampling during a spring situation. J. Plankt. Res. 2010 doi: 10.1093

Vohmann, A., Borcherding, J., Kureck, A., Bij de Vaate, A., Arndt, H. & Weitere, M. (2010): Strong body mass decrease of the invasive clam Corbicula fluminea during summer. Biological Invasions 12: 53-64

Wylezich C., Nies G., Mylnikov A. P., Tautz D. & Arndt H. (2010): An evaluation of the use of LSU rRNA D1-D5 region for DNA based taxonomy of eukaryotic protists. Protist 161: 342-352


Bergfeld, T., Scherwass, A., Ackermann, B., Arndt, H., Schöl, A. (2009): Comparison of the components of the planktonic food web in three large rivers (Rhine, Moselle and Saar)". River Res. Appl. 25 (10): 1232-1250

Kathol, M., Norf, H., Arndt, H., Weitere, M. (2009): Effects on temperature increase on the grazing of planctonic bacteria by biofilm-dwelling consumers. Aquat. Microb. Ecol. 55: 65-79

Loquay N., Wylezich C. & Arndt H. (2009): Composition of groundwater nanoprotist communities in different aquifers based on aliquot cultivation and genotype assessment of cercomonads. Fund. Appl. Limnol. 174 (3): 261-269

Norf, H., Arndt, H., Weitere, M. (2009): Effects of resource supplements on mature ciliate biofilms: an empirical test using a new type of flow cell. Biofouling 25 (6): 769-778

Norf, H., Arndt, H., Weitere, M. (2009): Responses of biofilm-dwelling ciliate communities to planctonic and benthic resource enrichment. Microb. Ecol. 57: 678-700

Risse-Buhl, U., Scherwass, A., Schlüssel, A., Arndt, H., Kröwer, S., Küsel, K. (2009): Detachment and motility of surface-associated ciliates at increased flow velocities. Aquat. Microb. Ecol. 55: 209-218

Vohmann, A., Mutz, M., Arndt, H., Weitere, M. (2009): Grazing impact and phenology of the freshwater sponge Ephydatia muelleri and the bryozoans Plumatella emarginata and Fredericella sultana under experimental warming. Fresw. Biol. 54: 1078-1092

Weitere, M., Vohmann, A., Schulz, N., Linn, C., Dietrich, D. Arndt, H. (2009): Linking environmental warming to the fitness of the invasive clam Corbicula fluminea. Global Change Biology 15: 2838-2851

Wey, J.K., Norf, H., Arndt, H., Weitere, M. (2009): Role of dispersal in shaping communities of ciliates and heterotrophic flagellates within riverine biofilms. Limnol. Oceanogr. 54(5): 1615-1626


Becks, L. and H. Arndt (2008): Transitions from stable equilibria to chaos, and back, in an experimental food web. Ecology 89 (11): 3222-3226

Nitsche, F. & Arndt, H. (2008).  A new choanoflagellate species from Taiwan: Morphological and molecular biological studies of Diplotheca elongata nov. spec. and D. costata. Europ. J. Protistol.44: 220-226

Weitere, M., Dahlmann, J., Viergutz, C., Arndt, H. (2008): Differential grazer-mediated effects of high summer temperatures on pico- and nanoplankton communities. Limnol. Oceanogr. 53 (2): 477-486

Wey, J.K., Scherwass, A., Norf, H., Arndt, H., Weitere, M. (2008): Effects of protozoan grazing within river biofilms under semi-natural conditions. Aquat. Microb. Ecol. 52: 283-296


Nitsche, F., Weitere M., Scheckenbach F., Hausmann K., Wylezich C., Arndt H. (2007): Deep sea records of choanoflagellates with a description of two new species. Acta Protozool. 46 (2):99-106

Norf, H., Arndt, H., Weitere, M. (2007): Impact of local temperature increase on the early development of biofilm-associated ciliate communities. Oecologia 151 (2): 341-350

Viergutz, C., Kathol, M., Norf, H., Arndt, H., Weitere, M. (2007): Control of microbial communities by the macrofauna: a sensitive interaction in the context of extreme summer temperatures? Oecologia 151 (1): 115-124

Willkomm, M., Schlüssel, A., Reiz, E., Arndt, H. (2007): Effects of microcurrents in the boundary layer on the attachment of benthic heterotrophic nanoflagellates. Aquat. Microb. Ecol. 48: 169-174

Wylezich C., Mylnikov A. P., Weitere M. & Arndt H. (2007): Distribution and Phylogenetic Relationships of Freshwater Thaumatomonads with a Description of the New Species Thaumatomonas coloniensis n. sp. J. Eukaryot. Microbiol. 54(4): 347-357


Borcherding, J., Murawski, S., Arndt, H. (2006): Population ecology, vertical migration and feeding of the Ponto-Caspian invader Hemimysis anomala in a gravel-pit lake connected to the River Rhine. Freshw. Biol. 51 (12): 2376-2387

Hausmann, K., Selchow, P., Scheckenbach, F., Weitere, M., Arndt, H. (2006): Cryptic Species in a Morphospecies Complex of Heterotrophic Flagellates: the Case Study of Caecitellus spp. Acta Protozool. 45: 415-431

Scheckenbach, F., Wylezich, C., Mylnikov, AP., Weitere, M., Arndt, H. (2006): Molecular comparisons of freshwater and marine isolates of the same morphospecies of heterotrophic flagellates. Appl Environ. Microb. 72 (10): 6638-6643


Becks, L., Hilker, F., Malchow, H., Jürgens, K., Arndt, H.(2005): Experimental demonstration of chaos in a microbial food web. Nature 435: 1226-1229

Kreuzer, J., Auer, B., Scherwass, A., Arndt, H. (2005): Jahreszeitliche Sukzession und vertikale Verteilung des Zooplanktons im Fühlinger See. Ökologie und nachhaltige Entwicklung von Sport- und Freizeitseen 1: 77-94

Scherwass, A., Fischer, Y., Arndt, H.(2005): Detritus as a potential food source for protozoans: consumption and digestion of plant detritus by a heterotrophic flagellate (Chilomonas paramecium) and a ciliate (Tetrahymena pyriformis). Aquat. Ecol. 39(4): 439-445

Scherwass, A., Arndt, H. (2005): Structure, dynamics and control of the ciliate fauna in the potamoplankton of the river Rhine. Archiv Hydrobiol 164(3): 287-307

Stauffer, D., Arndt, H. (2005): Simulation and experiment of extinction or adaptation of biological species after temperature changes. Int. J. Mod. Phys. C. 16 (3): 389-392

Weitere, M., Scherwass, A., Sieben, K.-T., Friedrich, G., Arndt, H. (2005): Planctonic Food Web Structure and Potential Carbon Flow in the Lower River Rhine with a Focus on the Role of the Protozoans. River Res. Appl. 21: 535-549

Scheckenbach, F., Wylezich, C., Weitere, M., Hausmann, K., Arndt H. (2005): Molecular identity of strains of heterotrophic flagellates isolated from surface waters and deep-sea sediments of the South Atlantic based on SSU rDNA. Aquat. Microb. Ecol. 38(3):239-247


Auer,B., Czioska,E., Arndt,H. (2004): The pelagic community of a gravel pit lake: Significance of Coleps hirtus viridis (Prostomatida) and its role as a scavenger. Limnologica 34: 187-198

Auer, B., Elzer, U., Arndt, H. (2004): Comparison of pelagic food webs in lakes along a trophic gradient and with seasonal aspects: influence of resource and prdation. J. Plankt. Res. 26 (6): 697-709

Dietrich, D., Arndt, H. (2004): Benthic heterotrophic flagellates in an Antarctic melt water stream. Hydrobiologia 529: 59-70

Matz, C., Deines, P., Boenigk, J., Arndt, H., Eberl, L., Kjelleberg, S., Jürgens, K. (2004): Impact of Violacein-Producing Bacteria on Survival and Feeding of Bacterivorous Nanoflagellates. Aquat. Microb. Ecol. 70 (3): 1593-1599 


Arndt, H., Hausmann, K., Wolf, M. (2003): Deep-sea heterotrophic nanoflagellates of the Eastern Mediterranean Sea: qualitative and quantitative aspects of their pelagic and benthic occurence. Mar. Ecol. Progr. Ser. 256: 45-56

Arndt, H.,  Schmidt-Denter, K., Auer, B. & Weitere, M. (2003). Protozoans and biofilms. In: Fossil and Recent Biofilms (Krumbein, W.E., Paterson, D.M. & Zavarzin, G.A., Eds.) Kluwer Academic Publ., Dordrecht., p. 173-189

Auer, B., Kiertucki, E., Arndt, H. (2003): Distribution of heterotrophic nano- and microflagellates in sediments of two small mesotrophic lakes. Archiv Hydrobiol. 158: 127-144

Prast, M., Arndt, H., Schöl, A. (2003): Untersuchungen zum planktischen Nahrungsnetz in Rhein und Mosel mittels Videomikroskopie. Hydrobiologie und Wasserbewirtschaftung 47: 102-107

Weitere, M., Schmidt-Denter, K., Arndt, H. (2003): Laboratory experiments on the impact of biofilms on the plankton of a large river. Freshw. Biol. 48: 1983-1992

Weitere, M., Arndt, H. (2003): Structure of the heterotrophic flagellate community in the water column of the River Rhine (Germany). Europ. J. Protistol. 39: 287-300


Arndt, H., Borcherding, J., Kureck, A. (2002): The Ecology of Large Rivers – Preface. Int. Revue Hydrobiol. 87(2-3): 137

Boenigk, J. & Arndt, H. (2002) Bacterivory by heterotrophic flagellates: community structure and feeding strategies. Antonie van Leeuwenhoek 81: 465-480

Boenigk, J., Matz, C., Jürgens, K., Arndt, H. (2002): Food concentration-dependent regulation of food selectivity of interception-feeding bacterivorous nanoflagellates. Aquat. Microb. Ecol. 27: 195-202

Garstecki, T., Wickham, S.A., Arndt, H. (2002): Effects of experimental sediment resuspension on a coastal planktonic microbial food web. Estuarine, Coastal and Shelf Science 55: 751-762

Hausmann, K, Weitere, M., Wolf, M., Arndt, H. (2002): Meteora sporadica gen. nov. et sp. nov. (Protista incertae sedis) - an extraordinary free-living protist from the Mediterranean deep sea.  Europ. J. Protistol. 38:171-177
Matz, C., Boenigk, J. Arndt, H., Jürgens, K. (2002): Role of bacterial phenotypic traits in selective feeding of the heterotrophic nanoflagellate Spumella sp. Aquat. Microb. Ecol. 27: 137-148

Pfister, G., Auer, B., Arndt, H. (2002): Pelagic ciliates (Protozoa, Ciliophora) of different brackish and freshwater lakes – a community analysis at the species level.  Limnologica 32: 147-168

Scherwass, A., Wickham, St.A., Arndt, H. (2002): Determination of the abundance of ciliates in highly turbid running waters - an improved method tested for the River Rhine. Archiv Hydrobiol. 156: 135-143

Weitere, M., Arndt, H. (2002): Top-down effects on pelagic heterotrophic nanoflagellates (HNF) in a large river (River Rhine): Do losses to the benthos play a role? Freshw. Biol. 47: 1437-1450

Weitere, M., Arndt, H. (2002): Water discharge regulated bacteria/ heterotrophic nanoflagellates (HNF) interactions in the water column of the River Rhine. Microb. Ecol. 44: 19-29


Auer, B., Arndt, H. (2001): Taxonomic composition and biomass of heterotrophic flagellates in relation to lake trophy and season. Freshw. Biol. 46: 959-972

Boenigk, J., Matz, C., Jürgens, K., Arndt, H. (2001): Confusing selective feeding with differential digestion in bacterivorous nanoflagellates. J. Eukaryot. Microbiol. 48: 425-432

Boenigk, J., Matz, C., Jürgens, K., Arndt, H. (2001): The influence of preculture conditions and food quality on the ingestion and digestion process of interception feeding heterotrophic nanoflagellates. Microb. Ecol. 42: 168-176

Boenigk, J., Arndt, H., Cleven, E.-J. (2001): The problematic nature of fluorescently labeled bacteria (FLB) in Spumella feeeding experiments – an explanation by video microscopy. Archiv Hydrobiol. 152 (2): 329-338

Pfister, G., Auer, B., Arndt, H.(2001): Community analysis of pelagic ciliates in numerous different freshwater and brackish water habitats. Verh. Int. Verein. theor. angew. Limnol. 27: 3404-3408

Scherwass, A., Eimer, A., Arndt, H.(2001): Selective influence of filter-feeding benthic bivalves (Corbicula sp., Mytilus sp.) on planktonic ciliates. Verh. Int. Verein. theor. angew. Limnol. 27: 3315-3318 


Arndt, H., Dietrich, D., Auer, B., Cleven, E.-J., Gräfenhan, T., Weitere, M., & Mylnikov, A.P.(2000). Functional Diversity of Heterotrophic Flagellates in Aquatic Ecosystems. In: The Flagellates (Leadbeater, B.S.C. & Green, J.C., Eds.) Taylor & Francis Ltd, London. 240-268.

Boenigk, J., Arndt, H. (2000): Particle handling during interception feeding by four species of heterotrophic nanoflagellate. J. Eukaryot. Microbiol. 47: 350-358

Boenigk, J., Arndt, H. (2000):. Comparative studies on the feeding behaviour of two heterotrophic nanoflagellates: the filter-feeding choanoflagellate Monosiga ovata and the raptorial-feeding kinetoplastid Rhynchomonas nasuta. Aquat. Microb. Ecol. 22: 243-249

Dietrich, D., Arndt, H. (2000): Biomass partitioning of benthic microbes in a Baltic inlet: Relationships  between bacteria, algae, heterotrophic flagellates and ciliates. Mar. Biol. 136: 309-322

Garstecki, T., Arndt, H. (2000): Seasonal abundances and community structure of benthic rhizopods in  shallow lagoons of the southern Baltic Sea. Europ. J. Protistol. 36: 103-115

Garstecki, T., Verhoeven, R., Wickham, S.A., Arndt, H.(2000): Benthic-pelagic coupling: a comparison  of benthic and planktonic heterotrophic protistan community structure in shallow estuaries of the Southern Baltic. Freshw. Biol. 45: 147-167

Premke, K., Arndt, H. (2000): Predation on heterotrophic flagellates by protists: Food selectivity determined using a live-staining technique. Archiv Hydrobiol. 150: 17-28


Dietrich, D., Arndt, H. (1998): Untersuchungen zum benthischen und pelagischen mikrobiellen Nahrungsgewebe der Potter Cove, King George Island, Antarktische Halbinsel, unter besonderer Berücksichtigung der heterotrophen Flagellaten und Ciliaten. Ber. Polarforsch. 277: 25-26

Pfister, G., Arndt, H. (1998): Food selectivity and feeding behaviour in omnivorous filter-feeding ciliates: A case study for Stylonychia. Europ. J. Protistol. 34: 446-457


Arndt, H. (1997): Spielen Protozoen bei der Manipulation pelagischer Nahrungsnetze eine Rolle? Laufener Seminararbeitr. 3/97: 77-88

Jürgens, K., Arndt, H, Zimmermann, H. (1997): Impact of metazoan and protozoan grazers on bacterial biomass distribution in microcosm experiments. Aquat. Microb. Ecol. 12: 131-138


Posch, T., Arndt, H. (1996): The uptake of sub-micrometre and µm-sized detrital particles by bacterivorous and omnivorous ciliates. Aquat. Microb. Ecol. 10: 45-53

Schiewer, U., Arndt, H., Heerkloss ,R., Jost, G., Walter, T., Wasmund, N. (1996): Experiments with controlled carbon flux variations in shallow brackish waters. Rostock. Meeresbiolog. Beitr. 4: 57-65


Mathes, J., Arndt, H. (1995): Annual cycle of protozooplankton (ciliates, flagellates and sarcodines) in relation to phyto- and metazooplankton in Lake Neumühler See (Mecklenburg, Germany). Arch. Hydrobiol. 134: 337-358

Arndt, H. & Berninger, U. (1995): Protists in aquatic food webs - complex interactions. In: G. BRUGEROLLE and J.-P. MIGNOT (eds.): Protistological Actualities, Clermont-Ferrand, France, p. 224-232


Arndt, H., (ed.)(1994): Protozooplankton Ecology - Methods and Seasonal Successions. Mar. Microb. Food Webs 8 (1 & 2): 1-375

Arndt, H. (1994): Protozoen als wesentliche Komponente pelagischer Ökosysteme von Seen. In: E. AESCHT (Hrsg.): Die Urtiere – eine verborgene Welt. Kataloge OÖ. Landesmuseums N.F. 71: 111-147
Salbrechter ,M.,  Arndt, H. (1994): The annual cycle of protozooplankton in the alpine, mesotrophic Lake Mondsee (Austria). Mar. Microb. Food Webs, 8: 217-234
Mathes, J., Arndt, H. (1994): Biomass and composition of protozooplankton in relation to lake trophy in north German lakes. Mar. Microb. Food Webs. 8: 357-375
Jürgens, K., Arndt, H., Rothaupt, K.O. (1994): Zooplankton-mediated changes of bacterial community structure. Microb. Ecol. 27: 27-42


Arndt, H. (1993): A critical review of the importance of rhizopods (naked and testate amoebae) and actinopods (Heliozoa) in lake plankton. Mar. Microb. Food Webs 7: 3-29

Arndt, H. (1993): Rotifers as predators on components of the microbial web (bacteria, heterotrophic flagellates, ciliates) - a review. Hydrobiologia, 255/256: 231-246

Arndt, H., Krocker, M., Nixdorf, B., Köhler, A. (1993): Long-term annual and seasonal changes of meta- and protozooplankton in Lake Müggelsee (Berlin): effects of eutrophication, grazing activities, and the impact of predation. Int. Revue ges. Hydrobiol. 78: 379-402

Nixdorf, B., Arndt, H. (1993): Seasonal changes in the plankton dynamics of a eutrophic lake including the microbial web. Int. Revue ges. Hydrobiol. 78: 403-410


Arndt, H., Güde, H., Macek, M., Rothaupt, K.O. (1992): Chemostats used to model the microbial food web: evidence for the feedback effect of herbivorous metazoans. Arch. Hydrobiol. Beih. Ergebn. Limnol. 37: 187-194


Arndt, H. (1991): Population dynamics and production of estuarine planktonic rotifers in the southern Baltic: Brachionus quadridentatus (HERMANN, 1783). Acta ichthyologica et piscatoria 21: 7-15

Arndt, H. (1991): On the importance of protozooplankton in the eutrophication process of the Baltic. Int. Revue ges. Hydrobiol. 76: 387-396

Arndt, H., Mathes, J. (1991): Large heterotrophic flagellates form a significant part of protozooplankton biomass in lakes and rivers. Ophelia 33: 225-234
Arndt, H., Nixdorf, B. (1991): Spring clear-water phase in a eutrophic lake: control by herbivorous zooplankton enhanced by grazing on components of the microbial web. Verh. Internat. Verein. Limnol. 24: 879-883

Nixdorf, B., Arndt, H., Schierhorn, E. (1991): Shortterm response within the microbial food web to organic matter loading. Acta ichthyologica et piscatoria S21: 163-170


Arndt, H. (1990): Spielen Protozoen bei der Manipulation pelagischer Nahrungsnetze eine Rolle? Laufener Seminararbeitr. 3/97: 77-88

Arndt, H. (1990): Das pelagische "microbial web" in einem eutrophen Flachsee: Jahreszeitliche Unterschiede in der Wechselwirkung zwischen Proto- und Matazooplankton. Deutsche Gesellschaft für Limnologie, Erweiterte Zusammenfassung der Jahrestagung 1990: 112-116

Arndt, H., Jost, G. & Wasmund, N. (1990): Dynamics of pelagic ciliates in eutrophic estuarine waters: Importance of functional groups among ciliates and responses to bacterial and phytoplankton production. Arch. Hydrobiol. Beih. Ergebn. Limnol. 34: 239-24

Arndt, H., Schiewer, U., Jost, G., Wasmund, N., Walter, T., Heerkloss, R., Arlt, G., Arndt,E.-A. (1990): The importance of pelagic and benthic microfauna in a shallow-water community of the Darss-Zingst estuary, southern Baltic, during mesocosm experiments. Limnologica (Berlin) 20: 101-106

Arndt, H., Schröder, C., Schnese, W. (1990): Rotifers of the genus Synchaeta - an important component of the zooplankton in the coastal waters of the southern Baltic. Limnologica (Berlin) 21: 233-235

Schiewer, U., Arndt, H., Jost, G., Heerkloss, R. & Wasmund, N. (1990): Carbon flux dynamics in a shallow eutrophic estuary. Limnologica (Berlin) 20: 95-100



Arndt, H. (1989): Zooplankton production and its consumption by planktivores in a Baltic inlet. Proceedings of the 21st European Marine Biology Symposium, Gdansk, 14-19 September 1986, Polish Academy of Sciences - Institute of Oceanology, p. 205-214

Arndt,H., Heerkloss,R. (1989): Diurnal variation in feeding and assimilation rates of planktonic rotifers and its possible ecological significance. Int. Revue ges. Hydrobiol. 74: 261-272

Arndt, H., Waterstraat, A., Franke, D., Lenschow, U. (1989): Kurze Mitteilung über die Nahrung der kleinen Maräne (Coregonus albula L.) während eines Frühjahrsaspektes im Breiten Luzin (Mecklenburg). Fischeei-Forschung 27: 42-45

Arndt, H. (1989): Tageszeitliche Unterschiede in den Nahrungsketten pelagischer Ökosysteme. In: SCHÄLLER, G. & BÄHRMANN, R. (eds.)"Beiträge zur Ökophysiologie", Friedrich-Schiller-Universität Jena, p. 48-52


Arndt, H. (1988): Dynamics and production of a natural population of Brachionus plicatilis (Rotatoria, Monogononta) in a eutrophicated inner coastal water of the Baltic. Kieler Meeresforsch. Sonderh. 6: 147-153


Burckhardt, R., Arndt, H. (1987): Untersuchungen zur Konsumtion an Ciliaten durch Metazooplankter des Barther Boddens (südliche Ostsee). Meeersbiol. Beitr., WZ Rostock 36, math.-nat.R. 2: 22-26


Arndt, H. (1986): The relative importance of protozoans in a Baltic estuarine zooplankton community. Symp. Biol. Hung. 33: 257-261

Arndt, H., Schnese, W. (1986): Population dynamics and production of Acartia tonsa (Copepoda, Calanoida) in the Darss-Zingst estuary,  southern Baltic. Ophelia Suppl. 4: 329-334

Schiewer, U., Arndt, H., Baader, G., Börner, R., Evert, F.-K., Georgi,F., Heerkloss, R., Jost, G., Kell, V., Krüger, B., Walter, T.  (1986): The bounds and potential effects of NH4 (loading) on the pelagic system of a Baltic estuary. Limnologica (Berlin) 17: 7-28


Arndt, H. (1985): Eine Zählkammer für die mikroskopische Auswertung von Zooplanktonproben. WZ Rostock 34, math.-nat.R. 6: 30-31

Arndt, H., Kramer, H.-J., Heerkloss, R., Schröder, C. (1985): Zwei Methoden zur Bestimmung populationsdynamischer Parameter von Zooplanktern unter Laborbedingungen mit ersten Ergebnissen an Eurytemora affinis (Copepoda,Calanoida) und Synchaeta cecilia (Rotatoria, Monogononta). Meeresbiol. Beitr., WZ Rostock 34, math.-nat.R. 6: 17-21


Arndt, H., Debus ,L., Heerkloss, R., Schnese, W. (1984): Diurnal changes in the matter flux of a shallow-water ecosystem in a Baltic inlet. Ophelia Suppl. 3: 1-9

Arndt, H., Heerkloss, R., Schnese, W. (1984): Seasonal and spatial fluctuations of estuarine rotifers in a Baltic inlet. Limnologica 15: 377-385

Debus, L., Arndt, H. (1984): Nahrungsbiologische Untersuchungen an Jungfischpopulationen eines brackigen Flachwassergebietes des Barther Boddens (südliche Ostsee). Meeresbiol. Beitr., WZ Rostock 33, math.-nat.R. 6: 76-82

Heerkloss, R., Börner, R., Krummhaar, B., Arndt, H., Schnese, W. (1984): Growth inhibition in estuarine zooplankters due to the blue-green algae. Ophelia, Suppl. 3: 45-49

Heerkloss, R., Arndt, H., Hellwig, J., Vietinghoff, U., Georgi, F., WesselL, B., Schnese ,W. (1984): Consumption and assimilation by zooplankton related to primary production in the Baltic coastal inlet Barther Bodden. Limnologica 15: 387-394

Vietinghoff, U., Erdmann, N., Arndt, H., Kell, V., Hubert, M.-L. (1984): Integrated samples provide accurate means of parameters characterizing aquatic ecosystems. Int. Revue ges. Hydrobiol. 69: 121- 131

Vietinghoff, U., Hubert, M.-L., Heerkloss, R., Arndt, H., Schnese, W. (1984):  A mathematical submodel for the planktic Rotatoria in the ecosystem of the Barther Bodden. Int. Revue ges. Hydrobiol. 69: 159-172


Jansen, W., Arndt, H., Heerkloss, R. (1983): Die Rolle von Neomysis integer LEACH (Mysidacea, Crustacea) im Stoffumsatz des eutrophierten Brackwassergewässers Barther Bodden (südliche Ostsee). Wissenschaftliche Zeitschrift der Wilhelm-Pieck-Universität Rostock, Naturwissenschaftliche Reihe 5: 44-48


Arndt, H., Kobel, J., MIichalk, A., Wronna, F., Heerkloss, R.(1982): Untersuchungen zur Vertikalwanderung epibenthischer Copepoden mit Hilfe einer Planktonreuse. Meeresbiol. Beitr., WZ Rostock 31, math.-nat. R. 6: 57-60


Arndt, H., Schnese, W., Heerkloss, R.(1981): Untersuchungen zur räumlichen Verteilung des Zooplanktons in einem flachen inneren Küstengewässer der Ostsee. Meeresbiol. Beitr., WZ Rostock 30, math.-nat.R. 4/5: 43-48

Heerkloss, R., Arndt, H. (1981): Eine Lichtfalle zur Reinigung von Zooplanktonproben. Meeresbiol. Beitr., WZ Rostock 30, math.-nat.R. 4/5: 49-52


Heerkloss, R., Schnese, W., Arndt, H., Fischer, F. (1980): Konsumtionsrate und Vertikalwanderung des Zooplanktons in einem flachen inneren Küstengewässer der Ostsee. Meeresbiol. Beitr., WZ Rostock 29, math.-nat.R. 4/5: 73-76


Arndt, H., Borcherding, J., Kureck, A. (eds.) (2002): Ecological studies of Large Rivers. Int. Revue Hydrobiol.

Arndt, H. (ed.)(1994): Protozooplankton Ecology - Methods and Seasonal Successions. Marine Microbial  Food Webs 8(1) Gauthier-Villars, Montrouge Cedex, France.


Arndt, H.,  Schmidt-Denter, K., Auer, B., Weitere, M. (2003): Protozoans and biofilms. In: Fossil and Recent Biofilms (Krumbein, W.E., Paterson, D.M. & Zavarzin, G.A., Eds.) Kluwer   Academic Publ., Dordrecht., p. 173-189

Boenigk, J., Arndt, H. (2002): Bacterivory by heterotrophic flagellates: community structure and feeding strategies. Antonie van Leeuwenhoek 81: 465-480

Arndt, H., Dietrich, D., Auer, B., Cleven, E., Gräfenhan, T., Weitere, M., Mylnikov, A.P.(2000): Functional Diversity of Hetrotrophic Fflagellates in Aquatic Ecosystems. In: The Flagellates (Leadbeater, B.S.C. & Green, J.C., Eds.) Taylor & Francis Ltd, London. 240-268

Arndt, H., Jost, G., Wasmund, N. (1990): Dynamics of pelagic ciliates in eutrophic estuarine waters: Importance of functional groups among ciliates and responses to bacterial and phytoplankton production. Arch. Hydrobiol. Beih. Ergebn. Limnol. 34: 239-245

Schiewer, U., Arndt, H., Jost, G., Heerkloss, R., Wasmund, N. (1990): Carbon flux dynamics in a shallow eutrophic estuary. Limnologica (Berlin) 20: 95-100



Prof. Dr. H. Arndt

© Uni­ver­si­ty of Cologne
Prof. Dr. H. Arndt